##gff-version 3
##sequence-region C04SLm0101P13.1 1 91343
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match	444	990	1.000	-	0	ID=SGN-U328034_alignment_1;Target=SGN-U328034 1 464 +
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match_part	926	990	1.000	-	0	ID=GTH_SGN_U_tom_match_part_1;Target=SGN-U328034 1 65 +;Parent=SGN-U328034_alignment_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match_part	444	842	1.000	-	0	ID=GTH_SGN_U_tom_match_part_2;Target=SGN-U328034 66 464 +;Parent=SGN-U328034_alignment_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match	7737	10257	0.996	+	0	ID=SGN-U315799_alignment_1;Target=SGN-U315799 1 1565 -
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match_part	7737	9202	0.996	+	0	ID=GTH_SGN_U_tom_match_part_3;Target=SGN-U315799 1 1466 -;Parent=SGN-U315799_alignment_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match_part	9547	9574	0.500	+	0	ID=GTH_SGN_U_tom_match_part_4;Target=SGN-U315799 1467 1494 -;Parent=SGN-U315799_alignment_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match_part	9760	9771	0.583	+	0	ID=GTH_SGN_U_tom_match_part_5;Target=SGN-U315799 1495 1506 -;Parent=SGN-U315799_alignment_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match_part	9916	9949	0.618	+	0	ID=GTH_SGN_U_tom_match_part_6;Target=SGN-U315799 1507 1539 -;Parent=SGN-U315799_alignment_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match_part	10231	10257	0.630	+	0	ID=GTH_SGN_U_tom_match_part_7;Target=SGN-U315799 1540 1565 -;Parent=SGN-U315799_alignment_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match	10684	11512	1.000	+	0	ID=SGN-U328255_alignment_1;Target=SGN-U328255 1 829 +
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match_part	10684	11512	1.000	+	0	ID=GTH_SGN_U_tom_match_part_8;Target=SGN-U328255 1 829 +;Parent=SGN-U328255_alignment_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match	13395	14188	1.000	+	0	ID=SGN-U333200_alignment_1;Target=SGN-U333200 1 642 +
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match_part	13395	13541	1.000	+	0	ID=GTH_SGN_U_tom_match_part_9;Target=SGN-U333200 1 147 +;Parent=SGN-U333200_alignment_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match_part	13694	14188	1.000	+	0	ID=GTH_SGN_U_tom_match_part_10;Target=SGN-U333200 148 642 +;Parent=SGN-U333200_alignment_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match	14168	16762	1.000	+	0	ID=SGN-U321226_alignment_1;Target=SGN-U321226 1 1504 +
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match_part	14168	14197	1.000	+	0	ID=GTH_SGN_U_tom_match_part_11;Target=SGN-U321226 1 30 +;Parent=SGN-U321226_alignment_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match_part	14422	14919	1.000	+	0	ID=GTH_SGN_U_tom_match_part_12;Target=SGN-U321226 31 528 +;Parent=SGN-U321226_alignment_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match_part	15600	16100	1.000	+	0	ID=GTH_SGN_U_tom_match_part_13;Target=SGN-U321226 529 1029 +;Parent=SGN-U321226_alignment_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match_part	16288	16762	1.000	+	0	ID=GTH_SGN_U_tom_match_part_14;Target=SGN-U321226 1030 1504 +;Parent=SGN-U321226_alignment_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match	19527	20218	0.996	+	0	ID=SGN-U328736_alignment_1;Target=SGN-U328736 1 692 +
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match_part	19527	20218	0.996	+	0	ID=GTH_SGN_U_tom_match_part_15;Target=SGN-U328736 1 692 +;Parent=SGN-U328736_alignment_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match	28494	28785	0.945	-	0	ID=SGN-U336817_alignment_1;Target=SGN-U336817 1 292 +
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match_part	28494	28785	0.945	-	0	ID=GTH_SGN_U_tom_match_part_16;Target=SGN-U336817 1 292 +;Parent=SGN-U336817_alignment_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match	60317	62143	0.999	+	0	ID=SGN-U313174_alignment_1;Target=SGN-U313174 1 1226 +
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match_part	60317	60642	1.000	+	0	ID=GTH_SGN_U_tom_match_part_17;Target=SGN-U313174 1 326 +;Parent=SGN-U313174_alignment_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match_part	60738	61018	1.000	+	0	ID=GTH_SGN_U_tom_match_part_18;Target=SGN-U313174 327 607 +;Parent=SGN-U313174_alignment_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match_part	61525	62143	0.998	+	0	ID=GTH_SGN_U_tom_match_part_19;Target=SGN-U313174 608 1226 +;Parent=SGN-U313174_alignment_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match	60325	61604	1.000	+	0	ID=SGN-U313173_alignment_1;Target=SGN-U313173 1 573 +
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match_part	60325	60642	1.000	+	0	ID=GTH_SGN_U_tom_match_part_20;Target=SGN-U313173 1 318 +;Parent=SGN-U313173_alignment_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match_part	60738	60863	1.000	+	0	ID=GTH_SGN_U_tom_match_part_21;Target=SGN-U313173 319 444 +;Parent=SGN-U313173_alignment_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match_part	60970	61018	1.000	+	0	ID=GTH_SGN_U_tom_match_part_22;Target=SGN-U313173 445 493 +;Parent=SGN-U313173_alignment_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match_part	61525	61604	1.000	+	0	ID=GTH_SGN_U_tom_match_part_23;Target=SGN-U313173 494 573 +;Parent=SGN-U313173_alignment_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match	60884	61407	0.998	+	0	ID=SGN-U334195_alignment_1;Target=SGN-U334195 1 524 +
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match_part	60884	61407	0.998	+	0	ID=GTH_SGN_U_tom_match_part_24;Target=SGN-U334195 1 524 +;Parent=SGN-U334195_alignment_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match	63421	64881	1.000	-	0	ID=SGN-U323949_alignment_1;Target=SGN-U323949 1 736 +
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match_part	64629	64881	1.000	-	0	ID=GTH_SGN_U_tom_match_part_25;Target=SGN-U323949 1 253 +;Parent=SGN-U323949_alignment_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match_part	64331	64454	1.000	-	0	ID=GTH_SGN_U_tom_match_part_26;Target=SGN-U323949 254 377 +;Parent=SGN-U323949_alignment_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match_part	63600	63949	1.000	-	0	ID=GTH_SGN_U_tom_match_part_27;Target=SGN-U323949 378 727 +;Parent=SGN-U323949_alignment_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match_part	63421	63429	0.778	-	0	ID=GTH_SGN_U_tom_match_part_28;Target=SGN-U323949 728 736 +;Parent=SGN-U323949_alignment_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match	66475	66972	1.000	-	0	ID=SGN-U327797_alignment_1;Target=SGN-U327797 1 498 +
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match_part	66475	66972	1.000	-	0	ID=GTH_SGN_U_tom_match_part_29;Target=SGN-U327797 1 498 +;Parent=SGN-U327797_alignment_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match	74565	75452	0.987	-	0	ID=SGN-U318079_alignment_1;Target=SGN-U318079 1 781 +
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match_part	74690	75452	0.987	-	0	ID=GTH_SGN_U_tom_match_part_30;Target=SGN-U318079 1 763 +;Parent=SGN-U318079_alignment_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match_part	74565	74582	0.778	-	0	ID=GTH_SGN_U_tom_match_part_31;Target=SGN-U318079 764 781 +;Parent=SGN-U318079_alignment_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match	74682	77465	0.991	-	0	ID=SGN-U318078_alignment_1;Target=SGN-U318078 1 1496 +
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match_part	77386	77465	1.000	-	0	ID=GTH_SGN_U_tom_match_part_32;Target=SGN-U318078 1 80 +;Parent=SGN-U318078_alignment_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match_part	77053	77135	1.000	-	0	ID=GTH_SGN_U_tom_match_part_33;Target=SGN-U318078 81 163 +;Parent=SGN-U318078_alignment_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match_part	74682	76021	0.990	-	0	ID=GTH_SGN_U_tom_match_part_34;Target=SGN-U318078 164 1496 +;Parent=SGN-U318078_alignment_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match	84697	85421	0.994	-	0	ID=SGN-U341941_alignment_1;Target=SGN-U341941 1 725 +
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	match_part	84697	85421	0.994	-	0	ID=GTH_SGN_U_tom_match_part_35;Target=SGN-U341941 1 725 +;Parent=SGN-U341941_alignment_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	gene	444	990	.	-	0	ID=GTH_SGN_U_tom_gene_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	mRNA	444	990	.	-	0	ID=GTH_SGN_U_tom_gene_1_AGS_1;Parent=GTH_SGN_U_tom_gene_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	exon	926	990	1.000	-	0	ID=GTH_SGN_U_tom_exon_1;Parent=GTH_SGN_U_tom_gene_1_AGS_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	exon	444	842	1.000	-	0	ID=GTH_SGN_U_tom_exon_2;Parent=GTH_SGN_U_tom_gene_1_AGS_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	gene	7737	10257	.	+	0	ID=GTH_SGN_U_tom_gene_2
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	mRNA	7737	10257	.	+	0	ID=GTH_SGN_U_tom_gene_2_AGS_1;Parent=GTH_SGN_U_tom_gene_2
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	exon	7737	9202	0.996	+	0	ID=GTH_SGN_U_tom_exon_3;Parent=GTH_SGN_U_tom_gene_2_AGS_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	exon	9547	9574	0.500	+	0	ID=GTH_SGN_U_tom_exon_4;Parent=GTH_SGN_U_tom_gene_2_AGS_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	exon	9760	9771	0.583	+	0	ID=GTH_SGN_U_tom_exon_5;Parent=GTH_SGN_U_tom_gene_2_AGS_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	exon	9916	9949	0.618	+	0	ID=GTH_SGN_U_tom_exon_6;Parent=GTH_SGN_U_tom_gene_2_AGS_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	exon	10231	10257	0.630	+	0	ID=GTH_SGN_U_tom_exon_7;Parent=GTH_SGN_U_tom_gene_2_AGS_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	gene	10684	11512	.	+	0	ID=GTH_SGN_U_tom_gene_3
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	mRNA	10684	11512	.	+	0	ID=GTH_SGN_U_tom_gene_3_AGS_1;Parent=GTH_SGN_U_tom_gene_3
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	exon	10684	11512	1.000	+	0	ID=GTH_SGN_U_tom_exon_8;Parent=GTH_SGN_U_tom_gene_3_AGS_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	gene	13395	16762	.	+	0	ID=GTH_SGN_U_tom_gene_4
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	mRNA	13395	16762	.	+	0	ID=GTH_SGN_U_tom_gene_4_AGS_1;Parent=GTH_SGN_U_tom_gene_4
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	exon	13395	13541	1.000	+	0	ID=GTH_SGN_U_tom_exon_9;Parent=GTH_SGN_U_tom_gene_4_AGS_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	exon	13694	14197	1.000	+	0	ID=GTH_SGN_U_tom_exon_10;Parent=GTH_SGN_U_tom_gene_4_AGS_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	exon	14422	14919	1.000	+	0	ID=GTH_SGN_U_tom_exon_11;Parent=GTH_SGN_U_tom_gene_4_AGS_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	exon	15600	16100	1.000	+	0	ID=GTH_SGN_U_tom_exon_12;Parent=GTH_SGN_U_tom_gene_4_AGS_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	exon	16288	16762	1.000	+	0	ID=GTH_SGN_U_tom_exon_13;Parent=GTH_SGN_U_tom_gene_4_AGS_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	gene	19527	20218	.	+	0	ID=GTH_SGN_U_tom_gene_5
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	mRNA	19527	20218	.	+	0	ID=GTH_SGN_U_tom_gene_5_AGS_1;Parent=GTH_SGN_U_tom_gene_5
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	exon	19527	20218	0.996	+	0	ID=GTH_SGN_U_tom_exon_14;Parent=GTH_SGN_U_tom_gene_5_AGS_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	gene	28494	28785	.	-	0	ID=GTH_SGN_U_tom_gene_6
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	mRNA	28494	28785	.	-	0	ID=GTH_SGN_U_tom_gene_6_AGS_1;Parent=GTH_SGN_U_tom_gene_6
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	exon	28494	28785	0.945	-	0	ID=GTH_SGN_U_tom_exon_15;Parent=GTH_SGN_U_tom_gene_6_AGS_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	gene	60317	62143	.	+	0	ID=GTH_SGN_U_tom_gene_7
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	mRNA	60317	62143	.	+	0	ID=GTH_SGN_U_tom_gene_7_AGS_1;Parent=GTH_SGN_U_tom_gene_7
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	exon	60317	60642	1.000	+	0	ID=GTH_SGN_U_tom_exon_16;Parent=GTH_SGN_U_tom_gene_7_AGS_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	exon	60738	61018	1.000	+	0	ID=GTH_SGN_U_tom_exon_17;Parent=GTH_SGN_U_tom_gene_7_AGS_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	exon	61525	62143	0.998	+	0	ID=GTH_SGN_U_tom_exon_18;Parent=GTH_SGN_U_tom_gene_7_AGS_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	mRNA	60884	61407	.	+	0	ID=GTH_SGN_U_tom_gene_7_AGS_2;Parent=GTH_SGN_U_tom_gene_7
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	exon	60884	61407	0.998	+	0	ID=GTH_SGN_U_tom_exon_19;Parent=GTH_SGN_U_tom_gene_7_AGS_2
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	mRNA	60325	61604	.	+	0	ID=GTH_SGN_U_tom_gene_7_AGS_3;Parent=GTH_SGN_U_tom_gene_7
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	exon	60325	60642	1.000	+	0	ID=GTH_SGN_U_tom_exon_20;Parent=GTH_SGN_U_tom_gene_7_AGS_3
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	exon	60738	60863	1.000	+	0	ID=GTH_SGN_U_tom_exon_21;Parent=GTH_SGN_U_tom_gene_7_AGS_3
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	exon	60970	61018	1.000	+	0	ID=GTH_SGN_U_tom_exon_22;Parent=GTH_SGN_U_tom_gene_7_AGS_3
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	exon	61525	61604	1.000	+	0	ID=GTH_SGN_U_tom_exon_23;Parent=GTH_SGN_U_tom_gene_7_AGS_3
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	gene	63421	64881	.	-	0	ID=GTH_SGN_U_tom_gene_8
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	mRNA	63421	64881	.	-	0	ID=GTH_SGN_U_tom_gene_8_AGS_1;Parent=GTH_SGN_U_tom_gene_8
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	exon	64629	64881	1.000	-	0	ID=GTH_SGN_U_tom_exon_24;Parent=GTH_SGN_U_tom_gene_8_AGS_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	exon	64331	64454	1.000	-	0	ID=GTH_SGN_U_tom_exon_25;Parent=GTH_SGN_U_tom_gene_8_AGS_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	exon	63600	63949	1.000	-	0	ID=GTH_SGN_U_tom_exon_26;Parent=GTH_SGN_U_tom_gene_8_AGS_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	exon	63421	63429	0.778	-	0	ID=GTH_SGN_U_tom_exon_27;Parent=GTH_SGN_U_tom_gene_8_AGS_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	gene	66475	66972	.	-	0	ID=GTH_SGN_U_tom_gene_9
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	mRNA	66475	66972	.	-	0	ID=GTH_SGN_U_tom_gene_9_AGS_1;Parent=GTH_SGN_U_tom_gene_9
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	exon	66475	66972	1.000	-	0	ID=GTH_SGN_U_tom_exon_28;Parent=GTH_SGN_U_tom_gene_9_AGS_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	gene	74565	77465	.	-	0	ID=GTH_SGN_U_tom_gene_10
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	mRNA	74682	77465	.	-	0	ID=GTH_SGN_U_tom_gene_10_AGS_1;Parent=GTH_SGN_U_tom_gene_10
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	exon	77386	77465	1.000	-	0	ID=GTH_SGN_U_tom_exon_29;Parent=GTH_SGN_U_tom_gene_10_AGS_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	exon	77053	77135	1.000	-	0	ID=GTH_SGN_U_tom_exon_30;Parent=GTH_SGN_U_tom_gene_10_AGS_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	exon	74682	76021	0.990	-	0	ID=GTH_SGN_U_tom_exon_31;Parent=GTH_SGN_U_tom_gene_10_AGS_1
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	mRNA	74565	75452	.	-	0	ID=GTH_SGN_U_tom_gene_10_AGS_2;Parent=GTH_SGN_U_tom_gene_10
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	exon	74690	75452	0.987	-	0	ID=GTH_SGN_U_tom_exon_32;Parent=GTH_SGN_U_tom_gene_10_AGS_2
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	exon	74565	74582	0.778	-	0	ID=GTH_SGN_U_tom_exon_33;Parent=GTH_SGN_U_tom_gene_10_AGS_2
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	gene	84697	85421	.	-	0	ID=GTH_SGN_U_tom_gene_11
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	mRNA	84697	85421	.	-	0	ID=GTH_SGN_U_tom_gene_11_AGS_1;Parent=GTH_SGN_U_tom_gene_11
C04SLm0101P13.1	GenomeThreader_SGN_U_tomato	exon	84697	85421	0.994	-	0	ID=GTH_SGN_U_tom_exon_34;Parent=GTH_SGN_U_tom_gene_11_AGS_1